The Montredon local fauna yielded very rare bats, represented by damaged isolated teeth. Only a few documents are available for this period of the European Neogene. ln this poor state of knowledge, the material represents three undetermined species, a supposed molossid and two vespertilionids. 

A fossil fish fauna, based on 5533 otoliths, from the La Paillade locality at Montpellier is described and figured. The otolith-bearing marls correlate to mammal zone MN l (Aguilar, 1982), and thus represent the earliest Miocene. The fish fauna consists of 30 taxa belonging to 20 families. Two species are new: Dussumieria sittigi and Liza gaudanti. The predominant faunal element is the Lesueurigobius vicínalis-species complex, composing 73% of all investigated otoliths. The palaeoecological analysis reveals a marine to euryhaline fish fauna living under tropical to subtropical conditions in the transition zone littoral - sublittoral. Water depth probably was more than 10 m. The scarcity of pelagic físhes suggests that the habitat was either a sheltered bay and/or far away from the open sea. Furthermore, some genera represented in the La Paillade fish fauna presently live exclusively in the Indopacific realm. Their presence strongly supports a broad connection between the Indian Ocean, the Mediterranean, and the Paratethys Seas during the Early Miocene (Aquitanian). From a palaeobiogeographical point of view, faunal relationships were found between the La Paillade fish fauna and both the Paratethys fish fauna and the fish fauna from the deposits in the Upper Rhinegraben and the Mayence and Hanau Basins (Germany).



  

Cranial anatomy of a late Miocene rhizomyid, Brafhyrhizomys cf. B. pilgrimi, provides new evidence on the origin of the dorsal, round infraorbital foramen of living rhizomyines. Primitive rhizomyids retain a myomorphous keyhole foramen with a long ventral slit that retracts upward during the evolutionary history of the Rhizomyidae. The primitive condition of the elongated ventral slit is represented by Kanisamys sivalensis. Among later burrowers the foramen shows progressive dorsal migration, the ventral slit terminating midway up the snout in B.tertracharax and B. choristos ; well above the midline of the snout in Brachyrhizamys cf. B. pilgrimi. Apparently this transformation began earlier among Rhizomyinae than among Tachyoryctinae and continued to a more derived stage in rhizomyines. ln living Rhizomyx the ventral slit is absent and only a high round hole remains at the anterior end of the zygomatic arch. 

A systematic analysis of an hipparionine horse assemblage from Maragheh, Iran is made. A brief orientation to systematic philosophy and informal superspecific characterizations of some Old World hipparionines is given as a background to this work. A character state analysis of skulls is made, and has revealed five distinct species. A character state and stratigraphic trend analysis of isolated check tooth and postcranial remains, with known provenance, is also made. These two combined analyses reveal that the most resolute discrimination of hipparionine species and their evolutionary relationships occurs when multiple character complexes of associated skulls, maxillary and mandibular dentitions are made. When this is not possible, skulls have provided the best basis for discriminating species and their evolutionary relationships. Traditional characters of isolated cheek teeth and postcranial remains are shown here to offer limited information content for hipparionine phylogenetic systematics. The systematic portion of this study includes a comprehensive description of cranial and postcranial remains, and has further corroborated the distinction of five species which belong to at least three superspecific groups including: «Hipparion» geltyi sp. nov., Group 1; Hipparion prostylum (s. l.), and Hipparion campbelli sp. nov., Group 3; «Hipparíon» aff. moldavicum and «Hipparion» ?matthewi, Group 2. These species stratigraphic ranges and evolutionary relationships are also given here and argued to be important for establishing future hipparionine geochronologic correlations between a number of Eurasian late Miocene provinces. 

On the Track of Ice Age Mammals justifie pleinement son titre car l'auteur, Anthony J. Sutcliffe, apporte au lecteur faits et interprétations qui l'amèneront à s'intéresser encore plus au passé récent et à l'avenir de son environnement et à la question de l'impact de l'homme sur la nature. Après les chapitres qui présentent les temps glaciaires et les divers témoignages qui nous en sont parvenus, les cinquième et sixième apportent les informations nécessaires à la compréhension des résultats que nous donnent les chercheurs: principes, moyens d'étude et limites des méthodes, difficultés de l'intégration des données dans un cadre stratigraphique, variabilité des signaux climatiques, variabilité de leur intensité selon l'endroit par rapport au centre de la glaciation, complications liées à la qualité inégale de l'enregistrement géologique, en mer et sur le continent. 

Fourteen distinct phyletical lineages which belong at least in three families: Ischyromyidae ALSTON, 1876, Gliridae THOMAS, 1896 and Theridomyidae ALSTON, 1876, have been identified after the study of more than 3600 rodent dental remains from about twenty Early and Middle Eocene european localities. A systematical and phylogenetical revision of these rodents has been achieved. Nearly all the specific and generic diagnosis are emended. Several new combinations and synonymies are proposed. Four new species and two new genera, Euromys nov. (Ailuravinae) and Hartenbergeromys nov. (Microparamyini), are named and described. Euromys nov. gen. is known by three distinctive ypresian (MP 7 to MP 10 european reference levels) chronospecies. This new lineage is thought to be the direct ancestor of Meldimys MICHAUX, 1968 and Ailuravus RUTIMEYER, 1891. A new species of the genus Plesiarctomys BRAVARD, 1850, Pl. lapicidinarum from Condé-en-Brie (MP 8-9 reference level), allows to relate the Plesiarctomys lineage to the Pseudoparamys MICHAUX, 1964 one. The taxa Sparnacomys HARTENBERGER, 1971, Pantrogna HARTENBERGER, 1971, and Corbarimys MARANDAT, 1989 are erected to genus rank; the last one is not thought to be an Ischyromyidae. A new chronospecies of Pantrogna, P. marandati nov. sp. from the locality of Prémontré (MP 10 reference level), is described. This lineage is at the origin of two others, namely Masillamys TOBIEN, 1954, including M. mattaueri (HARTENBERGER, 1975) nov. comb. (MP 10 reference level), and Hartenbergeromys nov. gen., known from MP 10 (H. hautefeuillei nov. sp.) and MP 11 (H. parvus TOBIEN, 1954) reference levels. The phylogenetical position of Hartenbergeromys nov. gen., at the origin of the european family Theridomyidae, is discussed. The systematical and phylogenetical status of two probable Paramyinae, "Paramys" woodi MICHAUX, 1964 and an unnamed genus and species, are discussed. New populations of the primitive Gliridae Eogliravus HARTENBERGER, 1971 and of the primitive Theridomyidae Protadelomys HARTENBERGER, 1968, are described and assigned to previously known species.

  

The analysis of oxygene isotope variations as well as paleobotanical data suggest that the Oligocene/Miocene boundary corresponds to a transitional period marked by floristical and climatic variations. During this period, the pyreneo-alpine tectonics has contribued to modify the geography and western Europe landscapes. Faunal changes (appearances, extinctions, migrations) are observed in different mammalian groups, notably in the rodents. A study of the evolutionary trends and patterns in paleogene rodents is involved for the period ranging from level MP 28 of the Late Oligocene to the Early Miocene, including the Oligo-Miocene boundary.
The Rodents fauna from the sites of Venelles (Bouches-du-Rhône District, France) and Thezels (Lot, France), previously mentionned in litterature, have been studied. The first description of the Eomyidae of La Milloque (MP 29) has been completed. These faunas are compared to those from various localities dating from the considered period. In La Milloque, a new representative of the Eomys species is described next to a form close to Rhodanomys hugueneyae ENGESSER, 1987. It is the Eomys milloquensis nov. sp., the likely descendant of Eomys quercyi COMTE & VIANEY-LIAUD, 1987. Two new species are also described in Thezels: Eucricetodon thezelensis nov. sp., resulting from a likely and local evolution of Eucricetodon praecursor (SCHAUB, 1925) from La Milloque, which, in the same geographic area, could be at the origin of Eucricetodon hesperius ENGESSER, 1985 from Paulhiac. Plesiosminthus admyarion nov. sp., quite distinct from Plesiosminthus schaubi VIRET, 1926, which announces Plesiosminthus myarion SCHAUB 1930. Venelles 'Plesiosminthus schaubi population is considered as a sub-species, named Plesiosminthus schaubi meridionalis nov. subsp. New phylogenetic patterns are proposed. Among the Eomyidae, a quantification of various features of the M1-2/ crown (hypsodonty, degree of abrasion, occlusal angle, state of development of the I and V anticlines), and a comparison with the occlusal diagram of the other teeth among various other populations allows a more efficient separation of Eomys and Rhodanomys genera. In Western Europe, and within this period, it finally does not seem possible to gradually connect the genus Eomys to the genus Rhodanomys. The evolution of the Eomys quercyi - milloquensis lineage seems to underline a similar evolution to that which may have led from the Eomys to the Rhodanomys form. The latter which appears totally accomplished at level MP 29 of the Oligocene is considered as an immigrant. If we compare the most representative species of the Venelles, Thezels, and Coderet sites, (i.e. Rhodanomys, Eucricetodon, Adelomyarion, Peridyromys, Plesíosminthus), it becomes impossible to confirm their biochronological separation. The noticeable differences between the populations may be interpreted as geographical variations. An explanation to these variations, and to fauna's evolution during the Late Oligocene and Early Miocene can be found in the environmental modifications, supported by isotopic, paleobotanical and sedimentologic analysis. A tentative reconstruction of the environments is attempted by the cenogram method. The analysis of the fluctuations of fauna's diversity shows variations which may be correlated to a drop in temperature at MP 29, during the Late Oligocene, followed by an increase in temperature along with an aridity phenomenom, during the basal Miocene (MN O).The confrontation of various methods give the opportunity of reconstituting and comparing the evolution of the environment of three sequences of sites chosen from different regions. Ecological affinities of various rodents' species are being examined. It is possible to consider that the integration of all the conclusions resulting from this study should lead to an explanation to the evolution of rodents for the period around the Oligocene-Miocene boundary. The site of Coderet- level 3- would be posterior to the latter, at the beginnig of the Miocene, and would mark the level MN 0 of the Aquitanian.

  

Fossils of the Trionychidae, Bataguridae or Emydidae, Palaeophis and Crocodylia from Chattahoochee, NW Florida, USA, represent the first report of an Eocene reptile fauna from Florida. 

The litoptem Proterotheriidae are extinct endemic South American ungulates frequently used as an example of evolutionary convergence with the horses. They were considered to be exclusively Tertiary representatives with the youngest record being in the late Pliocene, before the appearence of the equids and cervids during the Great American Interchange. Two undoubted Pleistocene records in Argentina and the specimen here described from Uruguay, confirm the persistence of the proterotherids into that period. In the Quaternary, these ungulates are found outside the typical pampean region and probably were confined to a few northern and warmer more forested relictual microhabitats.

  

Two new species, Omanodon minor n. g., n. sp. and Shizarodon dhofarensis n. g., n. sp., known from fifteen isolated teeth, are described here as the first adapiform primates (?Anchomomyini) recognizable to date in the Taqah material (early Oligocene of Sultanate of Oman).

Omanodon minor n. g., n. sp. displays special morphological similarity to the adapid tribe Anchomomyini from the Eocene of Europe, and especially to the Anchomomys lineage. Resemblances with the extant lemurifonn Microcebus are also noticeable and could be regarded as supporting Schwartz & Tattersall (1983) hypothesis of special relationships between the anchomomyine adapids and the cheirogaleid lemuriformes. However, these morphological affmities can be interpreted, altematively, as the results of parallelisms: important differences in upper molars indicate that the resemblances of cheirogaleids and Omanodon minor n. g., n. sp. are indeed probably due to parallelisms. Phyletic relationship of O. minor n. g., n. sp. to Anchomomyini is finally the most likely hypothesis.

Shizarodon dhofarensis n. g., n. sp., although much more poorly known, is closely related to Omanodon minor n. g., n. sp., at least at a familial level. The general morphology of this species suggests
also a close link with adapid Anchomomyini, although precise relationships within this tribe remain obscure. Interesting resemblances of Shizarodon dhofarensis n. g., n. sp. to Djebelemur martinezi lower molars (early Eocene of Tunisia) are also noticeable. These resemblances are even stronger than those betwen Omanodon minor and Djebelemur martinezi. However the very bunodont upper molars referred to D. martinezi are unusual for adapids, and there are moreover some notable differences in their lower molars. Thus resemblances in Djebelemur and Shizarodon are probably due to paralellisms.

Because of the fragmentary nature of the material and of possible parallelisms, the systematic position of Omanodon and Shizarodon within adapiformes cannot however yet be established definitively. 

L'ouvrage est divisé en trois grandes parties traitant successivement: I. l'anatomie ("Proboscidean flesh and bones"), en abordant la taxonomie, l'allure générale et l'éco-éthologie des proboscidiens actuels et fossiles; II. les causes de mortalité collective naturelle et provoquée chez les actuels; III. les documents fossiles et l'extinction des formes anciennes d'Amérique du Nord. 

The Causses" is a near 3400 km2 large plateau located in the south of France. Here the first dinosaur footprints where found in 1935. After this, this area has yielded an ever-increasing number of ichnites now in excess of 500 specimens. These latter, 15 to 50 cm long, tridactyl or tetradactyl footprints of generally biped animals, were discovered at the surface of Hettangian to lower Sinemurian dolomite layers within 4 distinct stratigraphic units. The 35 sites bearing ichnites are located on the plateau margin. For the first time, morphologic characters studied through descriptive statistic methods with the usual parameters and classical Student and Snédecor tests, allowed us, to divide the whole set of biped traces into 6 ichnospecies. Their definitions are further constrained by multivariate statistical results using Principal Component Analysis (PCA), Factor Analysis of correspondances (FAC) and Discriminant Analysis (DA). All have confirmed the morphologic observations. So that now, the following taxa are identified : Grallator variabilis, G. lescurei, G. sauclierensis, G. minusculus, Eubrontes giganteus, Dilophosauripus williamsi, cf. Moraesichnium, Orníthopus fabrei nov ichnosp. The more immediately visible differences relate to the interdigital II-IV divarication and the digit length ratio. To this panel, we must add Batrachopus deweyi and shapes suggesting Trisauropodichnus and/or Anomoepus. Among all ichnite associations described in the lower Liasic, the New England assemblage presents the most affinities with ours. It shows the ichnotaxa Grallator, Dilophosauripus, Eubrontes, Batrachopus without forgetting Ornithopus fabrei nov. ichnosp. which is close to Ornithopus gallinaceus from the Massachusetts and Connecticut basins. On comparing the present early Jurassic ichnofauna of the Causses with the ones of the Middle and Upper Triassic formations of the eastem border of the Massif Central (France), it appears that tridactyl footprints become more and more numerous and large from Triassic to Early Jurassic. In the Causses, these latest are prevalent but in Quercy (France), Poland, Italy, USA, they are also associated with Omithopoda, Thyreophora and Sauropoda ichnites. Footprint areas considered here were generaly under an arid climate. Animals that passed by were heavy and bulky possible Megalosaur trackmakers, and lighter and slender Coelophysids or Ceratosaurs. For all, these areas were pathways as the orientations of the trackways seem point out. The directions followed by these reptiles were without any important variation during the Hettango-Sinemurian stages. These areas were also used from time to time by Crocodilomorpha and may be tetradactyl (I-IV) bipedal avian Theropods. However, the number of such trackways in sites, sometimes substantial, should not lead us to overestimate the trackmakers populations. These last were probably relatively moderately abondant in this inter-supratidal swamp environment. In the Causses, ichnites are connected with former algo-laminated deposits (Algal mats) which were rapidly hardened by means of calcitisation of cyanobacteria. The result has been a moderate depth of footprints; autopodia disturbing only a few cm of the carbonate substrate. Other fossils have been discovered : invertebrates with thin bivalve and gastropod shells, crustaceans tests and plants. These latter suggest the existence of paleomangroves like environments but also continental vegetation periodically overruning the swamp environment during regression/transgression cycles. At these times, wooded parts of it, could become protecting, feeding, resting and nesting places.

  

Many types of insectivores have been described from specimens found in the Quercy phosphorites. These remains very often were not dated because they came from old collections. Recent excavations have permitted the situation of Amphidozotherium cayluxi FILHOL in the late Eocene. Two new genera are descrlbed based on material both from the old collections (Cryplotopos nov. g.) and from that recently recovered (Darbonetus nov. g., beginning of the middle Oligocene). Their systematic positions are revised and comparisons with American faunas are made. Amphidozotherium is not a talpid, but an erinaceoid belonging to an indeterminate family. Saturninia gracilis STHELIN is classified among the Nyctitheriinae, Myxomygale antiqua FILHOL among the Urotrichini Talpinae, and the genus Geotypus POMEL among the Scaptonichini Talpinae. A new study of the talpids from Auvergne has been rendered necessary. During the late Eocene and Oligocene precise morphology relationships existed between certain insectivores of Europe and North America. 

The object of this work is the study of the artiodactyls of the karstic locality of Lissieu near Lyon. Among the sixteen species counted, five (Hyperdichobune nobilis, Hyperdichobune sp., Haplabunodon mulleri, Tapirulus depereti and Pseudamphimeryx schlosseri) are cited for the first time in this fauna. The description of part of the anterior dentition of «Dichobune» langi permits the referal of this species to the genus Hyperdichobune. The new selenodont species Dichodon lugdunensis  described in this work belongs, with its ancestor Dichodon simplex, to a lineage distinct from that of Dichodan cartieri and Dichodon ruetimeyeri. Finally, the taxonomic reality of the Lissieu species, Lophiobunodon rhodanicum Deperet, is justified. All the other species have been compared with those of Egerkingen. Even though with most of them no difference in age can be shown between the two deposits, some however indicate clearly by their degree of evolution that Lissieu is slightly more recent than Egerkingen. ln the history of Eocene mammals, the Lissieu fauna, because of its homogeneity, can be considered as a sure reference fauna of the Auversian period. 

An increasing fossil record of dinosaur eggs and eggshells allows putting ootaxa within a chronostratigraphic framework, in order to study their distribution pattern leading eventually to their use as biochronological markers. For these purposes, high-quality data exists in four major regions; North America, South America, Europe and Asia (Central Asia and India). Most of the highly diverse dinosaur egg record has been dated as Latest Cretaceous in age (Campanian-Maastrichtian), reaching the Cretaceous-Tertiary boundary closer than the dinosaur bone record. However, dating continental sections is problematic and need to be carefully verified, as it appears when comparing the European dinosaur eggshell record from two well-studied areas. Ootaxa distribution in both sides of the Pyrenees (Tremp and Aix basins) shows comparable oospecies successions, but different chronology. This disagreement probably indicates that one or both successions have a wrong chronostratigraphic calibration.  

Le premier Symposium International sur les ceufs de dinosaures et leurs petits a connu un francs succès, à Isona, en Catalogne (Espagne) en 1999. I1 faisait suite à la publication en 1994 d'un premier ouvrage "Dinosaurs eggs and babies" édité par K. Carpenter, K. Hirsch et J. Homer. Entre 1994 et 1999, les nouvelles découvertes ont augmenté significativement, notamment celles d'ceufs embryonnés, et le nombre de chercheurs impliqués dans ce domaine a accompagné cet accroissement. Jusque là, l'étude de ces objets, les coquilles d'ceufs, est restée longtemps marginale, faute d'une méthodologie scientifique appropriée. 

A new Tertiary megadermatid is described from Rackham's Roost Site, a Pliocene limestone cave deposit on Riversleigh Station, northwestern Queensland, Australia. It appears to represent the first Australian record of Megaderma GEOFFROY, 1810, a genus otherwise known from Tertiary African and European taxa and the living Asian species M. spasma (LINNAEUS, 1758) and M. (Lyroderma) lyra PETERS, 1872. Megademza richardsi n. sp. is one of the smallest megademiatids known. It exhibits a mixture of plesiomorphic and autapomorphic features, the latter appearing to exclude it from being ancestral to any living megadermatid. The new species is one of eight megadermatids identified from the Australian fossil record, most of which are referable to Macroderma MILLER, 1906. 

Paleosols and the taphonomy of vertebrate fossils in the Eocene Kuldana Formation of northern Pakistan provide important information on the preservation and time-averaging of fossil assemblages. Morphologic, mineralogic, and chemical data as well as comparisons with Quaternary soils suggest that Kuldana paleosols formed under generally dry and oxidizing conditions over time intervals of less than 100 000 years and perhaps as short as 1000 years. The distribution of carbonate in Kuldana paleosols further indicates that the upper half of the profiles were acidic whereas the lower halves were alkaline. Vertebrate fossils are rare in Kuldana paleosols and occur primarily in well-cemented sandstones and conglomerates with abundant micritic and iron-stained nodules that were reworked from floodplain soils. The scarcity of vertebrate remains in Kuldana paleosols probably reflects a combination of acidic, dry, and oxidizing conditions in the upper half of the profiles and rapid floodplain sedimentation. Comparisons between the taphonomic characteristics of Kuldana channel fossil assemblages and bone accumulations in modem rivers provide a basis for estimating the length of time represented by Kuldana fossils from several important localities. Vertebrate fossil assemblages from Barbora Banda are characterized by a low-diversity paleofauna, partially articulated skeletons, and bones that are sorted by size and shape. Comparison with bone accumulations in modern rivers suggests that the fossils from Barbora Banda accumulated in 1 to 10 years. Vertebrate fossils from the Lower Kuldana in the Kala Chitta Hills region, typified by locality H-GSP 62, are characterized by a high-diversity paleofauna and generally random and unsorted fossil bone distributions, which suggest that the fossils from these localities represent longer time intervals than the Barbora Banda fossils. Based on the time estimates for Kuldana paleosol development, fossil assemblages in Kuldana channel deposits in the Kala Chitta Hills region probably represent time intervals of about 1000 years. 

In this article is described a new species of Nyctitheriidae with primitive characters: Leptacodon nascimentoi n. sp. from the early Eocene of Silveirinha (Portugal). 

A partial skull of Ardynomys russelli sp. nov. (Rodentia, Cylindrodontidae) is described. This was collected in the late Eocene of Alag Tsab locality in the eastem Gobi Desert, Mongolia. Ardynomys russelli sp. nov. is characterized by small size, brachyodont molars, and retention of P3. It represents the earliest record of the genus Ardynomys MATTHEW & GRANGER, 1925, in Asia.